Biology Department
Nicole Theodosiou, associate professor of Biological Sciences

Nicole Theodosiou

Job Title
Associate Professor of Biological Sciences
Co-Director of Biochemistry
Integrated Science and Engineering Complex (ISEC) 313-A
Biology, Biochemistry

Research interests

Evolution and development of the digestive tract in the vertebrate lineage. I study patterning (how different organs are ordered) and morphogenesis (how organs take on their final shape and function) of the digestive tract in vertebrate embryos and how the digestive tract evolved in the vertebrate lineage over 450 million years.

Teaching interests

Cellular Foundations of Life (BIO 104), Illustrated Organism (BIO 345), Developmental Biology (BIO 354), Topics in Organismal, Physiological Biology (BIO 488) and Australia Term Abroad (TAB)


  • Theodosiou, N. A. (2012). RNA in situ hybridization in whole mount embryos and cell histology adapted for marine elasmobranchs. JoVE, in press.
  • Theodosiou, N. A. and Parton, A. (2012). Establishing primary cultures of embryonic intestinal cells from the elasmobranch, Leucoraja erinacea. In Vitro Cellular and Developmental Biology – Animal, 48(7):413-417. DOI: 10.1007/s11626-012-9534-8.
  • Theodosiou, N. A. and *Simeone, A. (2012). Evidence of a rudimentary colon in the elasmobranch, Leucoraja erinacea. Development, Genes and Evolution, 222(4):237-43. DOI: 10.1007/s00427-012-0406-8.
  • *Simeone, A. and Theodosiou, N. A. (2010). Measuring water absorption in the digestive tract of Leucoraja erinacea. The Bulletin MDI Biol Lab, 49:35-6.
  • Richards, E. K., *Simeone, A. and Theodosiou, N. A., (2009). Water absorption in the spiral intestine of Leucoraja erinacea. The Bulletin MDI Biol Lab, 48:88-9.
  • Theodosiou, N. A., *Hall, D. A. and Jowdrey, A. L. (2007). Expression of Hoxa13 and Hoxd13 in Raja erinacea suggest an evolutionary role in hindgut patterning. Journal of Experimental Zoology Part B, 308B(4):442-453.

(* indicates undergraduate co-authors)

Additional media

Academic credentials

B.A., Swarthmore College; Ph.D., Yale University